I have found my own answer... From a review of the MSI X99S Gaming 9 AC Mobo
http://www.guru3d.com/articles-pages/msi-x99s-gaming-9-ac-motherboard-review,5.html
PCIe Lanes
The one thing that people grumble about the most is the relatively small number of available PCIe lanes (16) for graphics cards. If you are using a setup with two or more graphics cards (SLI/Crossfire), the PCIe bandwidth is limited at x8:x8. Here again the performance difference is trivial as running today's fastest cards barely utilizes all that bandwidth, but two x16 and then room for another x8 obviously is much better.
The eight core Core i7 5960X Haswell-E has 40 lanes (PCIe Gen 3.0) available.
The six core Core i7 5930K Haswell-E has 40 lanes (PCIe Gen...