do as the first guy told you hes right
''(Note 3) For optimum performance, if only one PCI Express graphics card is to be installed, be sure to install it in the PCIEX16 slot.
http://www.gigabyte.com/products/product-page.aspx?pid=3901#sp
now AMD cards will run at x4 in a x-fire configuration so you could use the other slots but why if not doing x-fire --ou want the full x16 on a single / primary card
if you read the notes from your boards link I gave you see it steps down sharing of the slots bandwidth
bottom line is why limit the card and cut your slots bandwith ??
unlike a 990fx that has 2 slots at full x16 [that's all part of chipset limitations ]