Each printer seems to think that when a print job is sent that it, the printer, is the one to do the work. Something is being duplicated...
Three things to do:
1) Go to each printer and print out their respective configuration pages.
2) Go to the router's admin pages and determine if there is some listing or diagram of connected devices.
3) Work with some command line utilities:
arp -a via the command prompt may prove helpful. Likewise for netstat.
Powershell can be very helpful: E.g. Get-NetIPAdress
Reference example:
https://www.channelpronetwork.com/blog/entry/powershell-ip-commands
Verify that all the configuration settings and IP addresses are indeed as required and expected.
Next try "reverse engineering" as...